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The Duke and Duchess of Sussex announced the happy news during an otherwise serious and at times harrowing interview with Oprah Winfrey on CBS.
When Oprah quizzed them on the sex of their newborn, Harry and pregnant Meghan confirmed: "A girl".
"To have a boy then a girl - what more can you ask for?" Harry said, with a smile.
When asked how he felt about having a daughter on the way, Harry said: "Grateful, having any child, any one or a two would have been amazing, but to have a boy and then a girl . . . Now we've got our family and we've got four of us."
The couple went on to tell the TV host they wouldn't have any more children after their daughter's arrival.
"Two is it," Meghan said with a smile, later revealing she is due to give birth this summer.
It wasn't all so cheery during the chat, as other topics that were broached during the chat included shocking allegations of concern over Archie's skin colour and the fact that Prince Harry was cut off financially when he left his duties.
In the special, Oprah also spoke with Meghan about her marriage, motherhood and philanthropic work. Plus, the couple revealed they were having a little girl.
There was also a heartbreaking moment where Meghan discussed the vitriol she received from the media, and criticised the team working with the Royal Family for failing to put untrue headlines to bed.
The Duchess even heartbreakingly revealed she was left feeling suicidal as a result of her experiences as a royal, adding that if she hadn't left things could have taken a very dark turn.
Prince Harry later discussed his mother, Princess Diana. On how she was treated after leaving the Royal Family, the Prince candidly said: "For me I'm just really relieved and happy to be sitting here talking to you with my wife by my side.
"Because I can't imagine what it must have been like for her going through this process by herself all those years ago, because it's been unbelievably tough for the two of us but at least we had each other."
The interview is currently being broadcast in the US on CBS, and is set to be shown here in the UK on Monday at 9pm on ITV.
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